Vascular Plants of Williamson County

Houstonia pusilla [Rubiaceae]
tiny bluets, star–violet, small bluet, innocence, southern bluet, angel eyes, quaker ladies

Houstonia pusilla Schöpf, tiny bluets, star–violet, small bluet, innocence, southern bluet, angel eyes, quaker ladies.  Annual, slender–taprooted, not rosetted, 1–stemmed at base, ± erect with ascending branches in canopy, < 10 cm tall; shoots with only cauline leaves at 2—5 nodes, glabrous.

Stems

Stems 4–sided and 4–ridged, < 1 mm diameter, with 2 ridges decurrent from each leaf (4 per node), weak, ridge edges sometimes vesicular and aging reddish.

Leaves

Leaves opposite decussate, simple, petiolate and fused at node with stipules; stipules 2 per node, ± triangular, ca. 0.8 × 0.8 mm, nearly colorless, acuminate or 2–toothed at tip, with several orangish, conic glands on margins near petiole; petiole broadly channeled, < 1—5 mm long; blade ovate, 4—8 × 2—4.5 mm, tapered at base, entire, acute to obtuse at tip, inconspicuously veined with only the faint midrib visible, upper surface minutely bumpy, lower surface smooth.

Inflorescence

Inflorescence

Flower

Flower bisexual ± radial, 7—9 mm across; calyx deeply 4–lobed, scarcely fused at base, green, glabrous; lobes suberect, somewhat unequal, triangular to narrowly ovate, 2.5—3 × 0.7—0.9 mm, pair on broader side wider, on edges with enlarged bases; corolla 4–lobed, trumpet–shaped, glabrous; tube flared at base, ca. 2 mm long, white with 4 greenish veins; throat ± cylindric with 4 faint ridges, initially ca. 2.5 mm long and exserted from calyx after anthesis, flaring upper portion cuplike, to 1 mm long, violet to bluish on outer surface, inner surface reddish purple with yellowish starlike pattern pointing to sinuses; lobes initially erect then spreading, ovate to broadly elliptic, 4—5 × 2—2.3 mm, light violet to bluish, upper throat and lobes minutely papillate, lower surface paler, veins inconspicuous; stamens 4, attached at top of corolla tube, included; filaments slender, to 0.4 mm long, pale green; anthers dorsifixed, dithecal, 0.5—0.6 mm long, light yellow, longitudinally dehiscent; pollen light yellow; pistil 1; ovary inferior, strongly 2–lobed with 4 shallow grooved and compressed side–to–side, ca. 0.5 × 1 mm, green, glabrous, 2–chambered each chamber with several ovules; style 0.6—0.7 mm long, 2–branched, lower portion nearly colorless and branches pale green, the branches ascending and at level of anthers, stigmas short papillate.

Fruit

Fruit capsule, with 4 spreading, triangular calyx lobes, to 16–seeded, each half resembling a gecko head, ca. 2.5 × 4.5 × 2 mm.

Seed

Seed helmetlike, ca. 1 mm long.

A. C. Gibson